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Guidelines for the Selection of Biological Heritage Sites

6. Section 1: Habitat Guidelines

6.3 GRASSLAND(1)

Gr1 Areas of ancient semi-natural limestone and neutral grasslands over 0.5 hectare identified as one or more of the following NVC(2) types:

MG3 Anthoxanthum odoratum - Geranium sylvaticum

MG4 Alopecurus pratensis - Sanguisorba officinalis

MG8 Cynosurus cristatus - Caltha palustris

CG2 Festuca ovina - Avenula pratensis

CG6 Avenula pubescens

CG9 Sesleria albicans - Galium sterneri

CG10 Festuca ovina - Agrostis capillaris - Thymus praecox

Application

This guideline is to be applied only to sites which have been identified as referable to one or more of the NVC types shown above by a competent field surveyor with adequate training and experience in the National Vegetation Classification. Sown wildflower meadows and semi-natural grasslands of known recent origin are not eligible for inclusion.

Justification

The presence of these NVC types indicates grassland which, although often of agricultural origin, has not been subject to intensive production involving use of inorganic nitrogen or post-war ploughing and reseeding. Although MG8, for example, was probably widespread in former times, most of these grasslands will always have been of localised occurrence. All are now rare as a result of agricultural improvement over many years. They are rich in native plant species including many which are mainly restricted to them, and are irreplaceable in practice.

Those non-agricultural grasslands which are eligible for inclusion here are likely to be those which originated before the widespread loss of species-rich agricultural grasslands, and would have been colonised from such grasslands which formerly existed in the same locality.


Gr2 (a) Areas of semi-natural acid grassland (as defined by NVC(2)) over 0.5 hectare in Landscape Character Tracts A, B and C.

Application

This guideline may be applied to any area of established acidic grassland, irrespective of origin, provided it fulfils the criteria set down in the guideline.

Justification

Acidic grassland of any kind is a rare habitat in the coastal zone.


Gr2 (b) Representative examples of acid grassland (as defined by NVC(2)) over 0.5 hectare may be included outside Landscape Character Tracts A, B and C where they contribute to the habitat and species diversity of sites selected on the basis of other guidelines.

Application

This may be applied to areas of acidic grassland which form part of an upland vegetation mosaic but is usually restricted to those which are relatively species-rich, including such characteristic species as Polygala serpyllifolia, and Carex species such as C. pilulifera, C. nigra or C. binervis.

Justification

Upland acidic grassland in Lancashire is generally of rather low diversity in terms of flowering plants at least. Its conservation value is generally assumed to be relatively low but is not well-studied.

Some areas have been identified as Biological Heritage Sites on the basis of their breeding birds (see species guidelines). Otherwise existing information on this habitat is inadequate for the formulation of more definite selection guidelines of the kind which appear for limestone or neutral grasslands.


Gr3 Areas of old established semi-natural grassland over 0.5 hectare, including sites referable to the following NVC,(2) types, with 10 or more species in Table 1:

MG1 Arrhenatherum elatius

MG5 Cynosurus cristatus - Centaurea nigra

MG6 Lolium perenne - Cynosurus cristatus

MG9 Holcus lanatus - Deschampsia cespitosa

MG10 Holcus lanatus - Juncus effusus

Application

Whilst appropriate sites will, in the main, be referable to one or more of the NVC types listed above, this guideline may be applied to grassland of any NVC type, or where NVC type has not been determined, provided that the conditions in this and the next paragraph are fulfilled. Sites should meet one or both of the following criteria. Either they are ancient grasslands (estimated to be at least 100 years old), or are other old-established grasslands which are indistinguishable in practice from ancient grasslands both in terms of their species composition and structure. Sown wildflower meadows, developing semi-natural grasslands of known recent origin, and other grasslands which do not meet either of these criteria are not eligible for inclusion here.

Table 1 includes a representative selection of species which are characteristic of species-rich (low-fertility) grasslands. It includes species of neutral, limestone and flushed grasslands; experience in Lancashire has shown that good examples of these major semi-natural grassland types, either singly or in combination (as is frequently the case on the ground), can be identified using this table. The species concerned should be reasonably well-distributed over the whole or a significant part of the site. Sites which support 10 or more species in Table 1 but where a high proportion of those species are very rare or restricted to non-typical patches or the edges of the site would not normally be included.

Justification

Ancient species-rich semi-natural grasslands are an important part of Lancashires critical environmental capital which is difficult or impossible to replace once destroyed. Now uncommon, they are being lost faster than any other type of wildlife habitat. They are extremely vulnerable to agricultural improvement, since many species are lost when soil fertility is increased, and also to neglect. It is therefore important to identify those that remain and to encourage their conservation by means of schemes such as Countryside Stewardship as a matter of urgency.

Grassland

Each of the NVC types listed under this guideline can vary widely in its conservation value from one site to another. Even MG5 grassland, which is usually unimproved species-rich agricultural grassland, can occasionally include sites which are not species-rich and of doubtful conservation value. This guideline provides a means of determining which of the sites referable to these NVC types are of high conservation value. The guideline also takes account of the facts that firstly, some sites of nature conservation importance are mixtures of different NVC types, and secondly that many sites have not yet been surveyed using NVC.

Those non-agricultural grasslands which are eligible for inclusion here are likely to be those which originated before the widespread loss of species-rich agricultural grasslands, and would have been colonised from such grasslands which formerly existed in the same locality.


Table 1. Plant species of semi-natural grasslands (3)
(See Guideline Gr3)
Scientific Name Common Name
Achillea ptarmica Sneezewort
Ajuga reptans Bugle
Alchemilla spp Ladys-mantle
Anemone nemorosa Wood Anemone
Anthyllis vulneraria Kidney vetch
Botrychium lunaria Moonwort
Briza media Quaking-grass
Caltha palustris Marsh-marigold
Campanula rotundifolia Harebell
Carex caryophyllea Spring Sedge
Carex disticha Brown Sedge
Carex flacca Glaucous Sedge
Carex hostiana Tawny Sedge
Carex pallescens Pale Sedge
Carex panicea Carnation Sedge
Carex pulicaris Flea Sedge
Carex spicata Spiked Sedge
Centaurea nigra Common Knapweed
Cirsium heterophyllum Melancholy Thistle
Conopodium majus Pignut
Crepis paludosa Marsh Hawks-beard
Dactylorhiza spp. Marsh and Spotted-orchids
Danthonia decumbens Heath-grass
Equisetum sylvaticum Wood Horsetail
Euphrasia spp Eyebright
Filipendula ulmaria Meadowsweet
Galium sterneri Limestone Bedstraw
Galium verum Ladys Bedstraw
Genista tinctoria Dyers Greenweed
Geranium columbinum Long-stalked Cranes-bill
Geranium pratense Meadow Cranes-bill
Geranium sylvaticum Wood Cranes-bill
Geum rivale Water Avens
Gymnadenia conopsea Fragrant Orchid
Helianthemum nummularium Common Rock-rose
Helictotrichon pubescens Downy Oat-grass
Helictotrichon pratense Meadow Oat-grass
Hyacinthoides non-scripta Bluebell
Hypericum pulchrum Slender St. Johns-wort
Hypochoeris radicata Cats-ear
Knautia arvensis Field Scabious
Lathyrus linifolius Bitter-vetch
Lathyrus pratensis Meadow Vetchling
Leontodon autumnalis Autumn Hawkbit
Leontodon hispidus Rough Hawkbit
Leucanthemum vulgare Oxeye Daisy
Linum cartharticum Fairy Flax
Listera ovata Common Twayblade
Lotus corniculatus Common Birds-foot-trefoil
Lotus pedunculatus Greater Birds-foot-trefoil
Luzula campestris/multiflora Field/Heath Wood-rush
Lychnis flos-cuculi Ragged-Robin
Ophioglossum vulgatum Adders-tongue
Orchis mascula Early-purple Orchid
Parnassia palustris Grass-of-Parnassus
Pedicularis sylvatica Lousewort
Pilosella officinarum Mouse-ear Hawkweed
Pimpinella major Greater Burnet-saxifrage
Pimpinella saxifraga Burnet-saxifrage
Polygala spp. Milkwort
Polygonum bistorta Common Bistort
Primula farinose Birds-eye Primrose
Primula veris Cowslip
Prunella vulgaris Selfheal
Ranunculus bulbosus Bulbous Buttercup
Rhinanthus minor Yellow-rattle
Sanguisorba officinalis Great Burnet
Sanguisorba minor Salad Burnet
Saxifraga granulata Meadow Saxifrage
Saxifraga tridactylites Rue-leaved Saxifrage
Scabiosa columbaria Small Scabious
Serratula tinctoria Saw-wort
Sesleria caerulea Blue Moor-grass
Stachys officinalis Betony
Stellaria graminea Lesser Stitchwort
Succisa pratensis Devils-bit Scabious
Taraxacum sect. Erythrosperma Dandelion
Taraxacum sect. Spectabilia or Naevosa Dandelion
Thalictrum flavum Common Meadow-rue
Thymus polytrichus Wild Thyme
Trifolium medium Zigzag Clover
Trisetum flavescens Yellow Oat-grass
Trollius europaeus Globeflower
Valeriana dioica Marsh Valerian
Veronica officinalis Heath Speedwell

(1) Note that "marshy grasslands" identified on the Phase 1 Habitat Survey are not distinguished as such by the NVC, but classified under either neutral grassland or swamp/fen or mire.
(2) National Vegetation Classification, published as Rodwell, (1992).
(3) Scientific and common names are taken from Stace 2nd Ed (1997)

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